London - Joe Marler has come out of international retirement to be named in England's Rugby World Cup training squad, while Danny Cipriani has also been included in the 35-man roster that was announced on Thursday.
Harlequins prop Marler, 28, last featured for England against South Africa in June 2018 before retiring due to personal reasons, but he has now reversed that decision.
Gloucester's Willi Heinz has won the battle for inclusion at scrumhalf, with Saracens' Ben Spencer also included, but Wasps' Dan Robson misses out.
England boss Eddie Jones said: "I saw Joe during the Barbarians' week and he indicated that he had a desire to come back.
"We have since had a couple of chats and we have given him the opportunity to do that. Now it is up to him to show us what he has got.
"We can't wait to have the full squad in place and to have players competing for those (final World Cup) 31 places.
"It is a process of building the team up, building the cohesion, the tactical understanding of the team and our adaptability. We will need to get those things right to win the World Cup, and that is our aim."
Gloucester flyhalf Cipriani won the Premiership player of the year and players' player of the year awards and now has the chance to push for a final World Cup place.
Harlequins prop Kyle Sinckler has been included after a hamstring injury scare but Harlequins fullback Mike Brown has missed out, while Saracens' Alex Lozowski has also been overlooked.
The continued absence of Dylan Hartley due to knee injuries leaves the former captain unlikely to make the World Cup in Japan, which starts in September.
"World Cup selection is a four-year process," said Jones.
"We have looked at the squad carefully, tried to build up a squad that has enough experience, attitude and energy to be winners and I think we have that.
"In the process, we have left out some players who can feel unlucky. For those guys who have missed out, the door is never closed but they have just got to be ready, as you never know when the opportunity comes."
England must submit their final 31-man World Cup squad on September 2, and they start their campaign against Tonga in Sapporo on September 22.
England World Cup training squad
Danny Cipriani (Gloucester), Joe Cokanasiga (Bath), Elliot Daly (Saracens), Owen Farrell (Saracens), George Ford (Leicester), Piers Francis (Northampton) Willi Heinz (Gloucester), Jonathan Joseph (Bath), Jonny May (Leicester), Ruaridh McConnochie (Bath), Henry Slade (Exeter), Ben Spencer (Saracens), Ben Te'o (unattached), Manu Tuilagi (Leicester), Anthony Watson (Bath), Ben Youngs (Leicester)
Dan Cole (Leicester), Luke Cowan-Dickie (Exeter), Tom Curry (Sale), Charlie Ewels (Bath), Ellis Genge (Leicester), Jamie George (Saracens), Maro Itoje (Saracens), Joe Launchbury (Wasps), Courtney Lawes (Northampton), Lewis Ludlam (Northampton), Joe Marler (Harlequins), Ben Moon (Exeter), Brad Shields (Wasps), Kyle Sinckler (Harlequins), Jack Singleton (Saracens), Sam Underhill (Bath), Billy Vunipola (Saracens), Harry Williams (Exeter), Mark Wilson (Sale)
In camp for rehabilitation: George Kruis (Saracens), Jack Nowell (Exeter), Mako Vunipola (Saracens)