Cardiff - England winger Chris Ashton was a late withdrawal from the side to face Wales in Cardiff on Saturday.
Ashton turned his ankle before training on Thursday and failed a fitness test on the morning of the match.
Matt Banahan is promoted from the bench with Delon Armitage joining the replacements.
Teams
Wales: 15 James Hook, 14 George North, 13 Jamie Roberts, 12 Gavin Henson, 11 Shane Williams, 10 Rhys Priestland, 9 Mike Phillips, 8 Toby Faletau, 7 Sam Warburton (capt), 6 Dan Lydiate, 5 Alun Wyn Jones, 4 Luke Charteris, 3 Craig Mitchell, 2 Lloyd Burns, 1 Paul James
Substitutes: 16 Huw Bennett, 17 Ryan Bevington, 18 Josh Turnbull, 19 Justin Tipuric, 20 Tavis Knoyle, 21 Scott Williams, 22 Aled Brew
England: 15 Ben Foden, 14 Matt Banahan, 13 Mike Tindall (capt), 12 Shontayne Hape, 11 Mark Cueto; 10 Toby Flood, 9 Richard Wigglesworth; 8 Nick Easter, 7 Hendre Fourie, 6 Tom Wood, 5 Courtney Lawes, 4 Louis Deacon, 3 Dan Cole, 2 Steve Thompson, 1 Alex Corbisiero
Substitutes: 16 Lee Mears, 17 Matt Stevens, 18 Tom Palmer, 19 James Haskell, 20 Danny Care, 21 Charlie Hodgson, 22 Delon Armitage
Referee: Alain Rolland (IRL)