Four changes for Wallabies

London - Australia recalled right wing Digby Ioane to their side for the Cook Cup international against England at Twickenham on Saturday in one of four changes to the side thrashed 33-6 by France.

Ioane, who has missed the Wallabies last two Tests with a knee injury, replaced Pat McCabe, out with a neck problem.

Berrick Barnes is selected a fullback after coming off the bench against France in Paris last weekend.

The Wallabies were hammered in the scrum by France but England, who've enjoyed their fair share of set-piece success against Australia in recent years, will face a front row bolstered by the return of experienced tighthead prop Ben Alexander.

A wrist injury has prevented Alexander from playing since last month's 25-19 win over Argentina at Rosario, which sealed second place in the Rugby Championship.

Meanwhile Sitaleki Timani returns to the second row after missing the France match with a back problem.

Timani and Alexander replace Kane Douglas, ruled out with a knee injury, and Sekope Kepu respectively.

Australia had hoped to recall world-class flanker David Pocock for Saturday's match following a knee problem, only for a calf strain to rule him out of the Test instead.

The talented Michael Hooper continues at seven in his absence, with Kurtley Beale continuing at fly-half.

"There aren't too many challenges in Test rugby that are greater than playing England at Twickenham," said under-pressure Australia coach Robbie Deans.

"We are playing an England side that has been building up under its new leadership, and will see Saturday as the chance to lay down a marker for the future," the New Zealander added.

"They will be excited by the opportunities they saw out of our performance last weekend.

"But it goes both ways -- Saturday is a significant opportunity for us as a group as well.

"It's going to be a great occasion, and one that the whole group is looking forward to."

England: 15 Alex Goode, 14 Chris Ashton, 13 Manu Tuilagi, 12 Brad Barritt, 11 Charlie Sharples, 10 Toby Flood, 9 Danny Care, 8 Thomas Waldrom, 7 Chris Robshaw (captain), 6 Tom Johnson, 5 Geoff Parling, 4 Tom Palmer, 3 Dan Cole, 2 Tom Youngs, 1 Joe Marler

Substitutes: 16 David Paice, 17 David Wilson, 18 Mako Vunipola, 19 Joe Launchbury, 20 Tom Wood, 21 Ben Youngs, 22 Owen Farrell, 23 Mike Brown

Australia: 15 Berrick Barnes, 14 Digby Ioane, 13 Adam Ashley-Cooper, 12 Ben Tapuai, 11 Nick Cummins, 10 Kurtley Beale, 9 Nick Phipps, 8 Wycliff Palu, 7 Michael Hooper, 6 Dave Dennis, 5 Nathan Sharpe (captain), 4 Sitaleki Timani, 3 Ben Alexander, 2 Tatafu Polota Nau, 1 Benn Robinson

Substitutes: 16 Stephen Moore, 17 James Slipper, 18 Sekope Kepu, 19 Radike Samo, 20 Liam Gill, 21 Brett Sheehan, 22 Mike Harris, 23 Drew Mitchell