Johannesburg - England coach Stuart Lancaster has named his side to face the Springboks in the second Test at Coca-Cola Park in Johannesburg on Saturday. Kick-off is at 17:00.
Jonathan Joseph will start his first England international. The London Irish centre comes in at No 13 in place of Manu Tuilagi, who switches inside for Brad Barritt in a side showing five changes - two positional - from the one that started the first Test.
Toby Flood replaces Owen Farrell at flyhalf, Ben Foden moves to fullback for the injured Mike Brown and David Strettle comes on to the left wing.
The pack remains the same while there are changes on the replacements' bench, where Alex Corbisiero gets his first chance in the series, in place of Paul Doran Jones, while Alex Goode takes over from Toby Flood.
Lancaster said: "I am delighted for JJ. He has trained very well and showed against the Barbarians and in his short time on the field in the first Test that he is ready to make the step to international rugby. To have two 21-year-old centres is exciting and we are looking forward to seeing this combination in action.
"Owen has had a fantastic season and I am sure he will make a significant impact from the bench. However, Toby has waited patiently for his opportunity and the time is right to allow him the chance to demonstrate his experience and skills in what will be a very tough challenge.
15 Pat Lambie, 14 JP Pietersen, 13 Jean de Villiers (captain), 12 Frans Steyn, 11 Bryan Habana, 10 Morné Steyn, 9 Francois Hougaard, 8 Pierre Spies, 7 Willem Alberts, 6 Marcell Coetzee, 5 Juandré Kruger, 4 Eben Etzebeth, 3 Jannie du Plessis, 2 Bismarck du Plessis, 1 Beast Mtawarira
Substitutes: 16 Adriaan Strauss, 17 Werner Kruger , 18 Flip van der Merwe, 19 Keegan Daniel, 20 Ruan Pienaar, 21 Wynand Olivier, 22 Bjorn Basson
15 Ben Foden, 14 Chris Ashton, 13 Jonathan Joseph, 12 Manu Tuilagi, 11 David Strettle, 10 Toby Flood, 9 Ben Youngs, 8 Ben Morgan, 7 Chris Robshaw (captain), 6 Tom Johnson, 5 Geoff Parling, 4 Mouritz Botha, 3 Dan Cole, 2 Dylan Hartley, 1 Joe Marler
Substitutes: 16 Lee Mears, 17 Alex Corbisiero, 18 Tom Palmer, 19 Phil Dowson, 20 Lee Dickson, 21 Owen Farrell, 22 Alex Goode